Mr Russell said they simply did not have the scientific knowledge at that time to explain Mr Levick's accounts of what he termed necrophilia.
"What is happening there is not in any way analogous to necrophilia in the human context," Mr Russell said. "It is the males seeing the positioning that is causing them to have a sexual reaction.
"They are not distinguishing between live females who are awaiting congress in the colony, and dead penguins from the previous year which just happen to be in the same position."
